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Wednesday, May 29, 2024

This Is An Operating System: Surgical By Demand Also Equating A Hand Of God[A Three Legged Race] As Ewe Are Tied Equating Masking Tape Or Word Emasculation At word Eval to equate E word Vowel word U Word A words Shin: Word Pane: Hand of God Equated Horse, There Are Four Inches[Hinges]In That Hand Equating Knuckles To State Sandwich?l. This Equates Word Renew Without Balance


 
Add’1[n[NOW]]:  The basis of the da Vinci[Vitruvian Man]man to the bell must have an influence to the Masonic.  In subtraction; word dread equated word sinful as word sinne equated word bone[Plasma].  The word sinned equated word blood.

Balance

Cantore Arithmetic is able to equate at equations that the impossible is the possible waiting to happen:  Mapping!!

Word loneliness equated word location[s] as word lonely equated word body.  The word equals word Holy as equated frustration to equal word hollow.  All hollows eve to genesis may give Cantore Arithmetic the fig[figurine] to word debt as the floor for word dirt[dead[death[dusted[dam[bottom[foot[six]]]]]]] as mud rises to word clay[Pot].  In addition, word dead equated words bed so whom is in the room:  6ft under.  Number six[6] foot[ft[‘]] equated 60:  The symbol for “feet” is an apostrophe. The symbol for “inches” is a quotation mark or a ditto mark. Hence, a man who was six feet two inches tall would be 6′2″. Feet is condensed with a punctuation ' or ft.

And, word this equated word Computation!!  Cantore Arithmetic has completed it’s first pass as word eternal measured word at 1000.  Yrs is a dabber[Dabs].  This Is Flounder, This is Fall however Fall is in word Season leaving word Left As A leaf in A Table:  Reset man whom titled two left feet.  The halls of Victrola[tittle] to than the halls of Vahalla as the table is from the Era of word been[chair[cave]]

Too represention of to word rise from the feet would gain at crossed legs to equate crossing the word street and equation word zing.  These basis to word grave and word condition[horse] to say that near death is word grave as word death is word boxed.  The word ribbon is worded at word message to word up as posting trot[Skin equated Posting trots].  The word Post to word equals is word death a word gain[Product Safeway on Cabrillo] to Petrini as Frozen Orange Juice equated word blood at word gem[stone].

The word gnu[cantelope[Cantor[barrel[bum]]]] to the Antelope is word canter[cup[three-beets[glass]]]:  The Urik (Russian: Урик) is a river in Irkutsk Oblast, Russia. It is a right:  At word right word equated righteous.  Whereas spelled and punctuated Urik (river) equated word Stone.

What is the origin of two left feet?

Idioms and Phrases:  Be clumsy, as in I'll never get the hang of this dance; I've got two left feet . This expression conjures up an image of feet that are not symmetrical, as left and right are, therefore causing imbalance or stumbling. It was first recorded in 1915.


You searched for

"BODY" in the KJV Bible


152 Instances   -   Page 1 of 6   -   Sort by Book Order   -   Feedback

1 Corinthians 15:44chapter context similar meaning copy save
It is sown a natural body; it is raised a spiritual body. There is a natural body, and there is a spiritual body.


1 Corinthians 15:38chapter context similar meaning copy save
But God giveth it a body as it hath pleased him, and to every seed his own body.


1 Corinthians 12:12chapter context similar meaning copy save
For as the body is one, and hath many members, and all the members of that one body, being many, are one body: so also is Christ.


2 Corinthians 12:3chapter context similar meaning copy save
And I knew such a man, (whether in the body, or out of the body, I cannot tell: God knoweth;)


1 Corinthians 6:18chapter context similar meaning copy save
Flee fornication. Every sin that a man doeth is without the body; but he that committeth fornication sinneth against his own body.


Philippians 3:21chapter context similar meaning copy save
Who shall change our vile body, that it may be fashioned like unto his glorious body, according to the working whereby he is able even to subdue all things unto himself.


Matthew 27:58chapter context similar meaning copy save
He went to Pilate, and begged the body of Jesus. Then Pilate commanded the bodyto be delivered.


2 Corinthians 4:10chapter context similar meaning copy save
Always bearing about in the body the dying of the Lord Jesus, that the life also of Jesus might be made manifest in our body.


Luke 11:34chapter context similar meaning copy save
The light of the body is the eye: therefore when thine eye is single, thy whole bodyalso is full of light; but when thine eye is evil, thy body also is full of darkness.


Ephesians 4:16chapter context similar meaning copy save
From whom the whole body fitly joined together and compacted by that which every joint supplieth, according to the effectual working in the measure of every part, maketh increase of the body unto the edifying of itself in love.


Matthew 6:22chapter context similar meaning copy save
The light of the body is the eye: if therefore thine eye be single, thy whole bodyshall be full of light.


2 Corinthians 12:2chapter context similar meaning copy save
I knew a man in Christ above fourteen years ago, (whether in the body, I cannot tell; or whether out of the body, I cannot tell: God knoweth;) such an one caught up to the third heaven.


1 Corinthians 12:16chapter context similar meaning copy save
And if the ear shall say, Because I am not the eye, I am not of the body; is it therefore not of the body?


1 Corinthians 12:15chapter context similar meaning copy save
If the foot shall say, Because I am not the hand, I am not of the body; is it therefore not of the body?


John 19:38chapter context similar meaning copy save
And after this Joseph of Arimathaea, being a disciple of Jesus, but secretly for fear of the Jews, besought Pilate that he might take away the body of Jesus: and Pilate gave him leave. He came therefore, and took the body of Jesus.


Matthew 6:25chapter context similar meaning copy save
Therefore I say unto you, Take no thought for your life, what ye shall eat, or what ye shall drink; nor yet for your body, what ye shall put on. Is not the life more than meat, and the body than raiment?


Matthew 10:28chapter context similar meaning copy save
And fear not them which kill the body, but are not able to kill the soul: but rather fear him which is able to destroy both soul and body in hell.


1 Corinthians 7:4chapter context similar meaning copy save
The wife hath not power of her own body, but the husband: and likewise also the husband hath not power of his own body, but the wife.


1 Corinthians 6:13chapter context similar meaning copy save
Meats for the belly, and the belly for meats: but God shall destroy both it and them. Now the body is not for fornication, but for the Lord; and the Lord for the body.


John 2:21chapter context similar meaning copy save
But he spake of the temple of his body.


1 Corinthians 12:27chapter context similar meaning copy save
Now ye are the body of Christ, and members in particular.


1 Corinthians 12:14chapter context similar meaning copy save
For the body is not one member, but many.


Colossians 2:17chapter context similar meaning copy save
Which are a shadow of things to come; but the body is of Christ.


Luke 23:52chapter context similar meaning copy save
This man went unto Pilate, and begged the body of Jesus.


Matthew 27:59chapter context similar meaning copy save
And when Joseph had taken the body, he wrapped it in a clean linen cloth,


Romans 7:24chapter context similar meaning copy save
O wretched man that I am! who shall deliver me from the body of this death?


Numbers 19:11chapter context similar meaning copy save
He that toucheth the dead body of any man shall be unclean seven days.


Job 19:17chapter context similar meaning copy save
My breath is strange to my wife, though I intreated for the children's sake of mine own body.


Job 19:26chapter context similar meaning copy save
And though after my skin worms destroy this body, yet in my flesh shall I see God:


Romans 6:12chapter context similar meaning copy save
Let not sin therefore reign in your mortal body, that ye should obey it in the lusts thereof.


 


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Common dab

From Wikipedia, the free encyclopedia
Common dab
Scientific classificationEdit this classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Class:Actinopterygii
Order:Pleuronectiformes
Family:Pleuronectidae
Genus:Limanda
Species:
L. limanda
Binomial name
Limanda limanda
Common dab range.[2]
Synonyms
  • Pleuronectes limanda Linnaeus, 1758
  • Liopsetta limanda (Linnaeus, 1758)
  • Pleuronectes limandulaBonnaterre, 1788
  • Limanda vulgaris Gottsche 1835

The common dab (Limanda limanda) is an edible flatfish of the family Pleuronectidae. It is a demersal fish native to shallow seas around Northern Europe, in particular the North Sea, where it lives on sandy bottoms down to depths of about 100 metres (330 ft). It can reach 40 centimetres (16 in) in length and can weigh up to 1 kilogram (2.2 lb), though most specimens grow no longer than 20 centimetres (7.9 in).[3][4]

Taxonomy and nomenclature[edit]

The etymology of the name dab is unclear, but the modern English use seems to originate from the Middle English dabbe.[5] It is first recorded in the late 16th century.[6]

The common dab was first named Pleuronectes limanda by Carl Linnaeus in the 1758 10th edition of Systema Naturae. It has also been moved to other genera, including Liopsetta, and is now known as Limanda limanda.[7]

Identification[edit]

Common dab caught while fishing

The common dab has a similar appearance to both the plaice and the flounder, and similarly has both its eyes normally on the right-hand side of its body. The upper surface is usually pale brown in colour with scattered darker blotches and speckles, but does not have the orange spots typical of a plaice. They are distinguished from flounder by their translucent body. The pectoral fins may be orange. The lateral line is marked by a distinctive semi-circular curve above the pectoral fin. The dorsal and anal fins form a gently rounded curve round the margin of the body. The scales have rough posterior edges and this fish has no large bony projections. A typical size is in the range 25 to 40 cm (10 to 16 in).[4][8]

Diet[edit]

The common dab's diet consists of zoobenthos organisms such as marine wormsmolluscssand eelsamphipodscrustaceansechinoderms and small pieces of fish.[4]

Distribution[edit]

The common dab is a bottom dweller, found in coastal waters in the northeastern Atlantic Ocean. Its range extends from the Bay of Biscay to Iceland and the White Sea and includes the North Sea and the western part of the Baltic Sea.[9]

Commercial fishing[edit]

The dab is an abundant fish and until recently was mostly ignored as a commercial fish, with most dab only retained when they were caught as by-catch of other targeted species.[10][11] However, the declining numbers of other food fish such as cod and haddock has seen dab become an increasingly important commercial species.[12] They are now targeted by an increasing number of commercial vessels, especially in the North Sea. A number of high-profile celebrity chefs such as Jamie Oliver have attempted to get people to eat more dab in order to take the pressure off the species of commercial fish which are currently heavily exploited.[13]

References[edit]

  1. ^ Monroe, T.; Costa, M.; Nielsen, J.; Herrera, J.; de Sola, L. (2014). "Limanda limanda"IUCN Red List of Threatened Species2014: e.T18214863A45790133. doi:10.2305/IUCN.UK.2014-3.RLTS.T18214863A45790133.en. Retrieved 11 November 2021.
  2. ^ International Union for Conservation of Nature (IUCN) 2014. Limanda limanda. In: IUCN 2015. The IUCN Red List of Threatened Species. Version 2015.2. "The IUCN Red List of Threatened Species". Archived from the original on 2014-06-27. Retrieved 2011-01-24.. Downloaded on 23 July 2015.
  3. ^ "Limanda limanda"Fishbase. 15 January 2009. Archived from the original on 2 March 2006. Retrieved 2009-05-12.
  4. Jump up to: a b c Picton, B.E.; Morrow, C.C. (2005). "Limanda limanda"Encyclopaedia of Marine Life of Britain and Ireland. Habitas Online. Archived from the original on 2 August 2005. Retrieved 2009-04-28.
  5. ^ "dab"The American Heritage Dictionary of the English Language, Fourth EditionHoughton Mifflin Harcourt. 2006. Archived from the original on January 4, 2013. Retrieved 2009-05-12.
  6. ^ "dab"Random House Unabridged DictionaryRandom House. 2006. Archived from the original on January 4, 2013. Retrieved 2009-05-12.
  7. ^ "Synonyms of Limanda limanda"Fishbase. 7 May 2005. Retrieved 2009-05-12.
  8. ^ "Dab: Limanda limanda". NatureGate. Retrieved 2013-12-17.
  9. ^ "Species factsheet: Limanda limanda". Food and Agriculture Organization of the United Nations. Retrieved 2013-12-17.
  10. ^ Burton, Maurice; Burton, Robert (2002). "Dab". The international wildlife encyclopedia. Vol. 10. Marshall Cavendish. pp. 634–5. ISBN 978-0-7614-7266-7.
  11. ^ North Sea Task Force (1993). North Sea quality status report 1993. Fredensborg, Denmark: Olsen and Olsen. p. 70. ISBN 978-1-872349-07-7.
  12. ^ "Dab". British Sea Fishing. 6 September 2012. Retrieved 8 May 2014.
  13. ^ "Mediterranean-style Dab". JamieOliver.com. Retrieved 8 May 2014.

External links[edit]

Urik (river)

Urik
Physical characteristics
MouthBelaya
 • coordinates
52.9692°N 102.0007°E
Length210 km (130 mi)
Basin size3,520 km2 (1,360 sq mi)
Basin features
ProgressionBelaya→ Angara→ Yenisey→ Kara Sea

The Urik (RussianУрик) is a river in Irkutsk OblastRussia. It is a right tributary of the Belaya. It is 210 kilometres (130 mi) long, and has a drainage basin of 3,520 square kilometres (1,360 sq mi).[1]

References[edit]

External links[edit]

Wildebeest

This is a good article. Click here for more information.
From Wikipedia, the free encyclopedia
(Redirected from Gnu)
Wildebeest
Temporal range: 2.5–0 Ma Early Pleistocene – present
Blue wildebeestC. taurinus
Ngorongoro CraterTanzania
Black wildebeestC. gnou
Krugersdorp Game ReserveSouth Africa
Scientific classificationEdit this classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Class:Mammalia
Order:Artiodactyla
Family:Bovidae
Subfamily:Alcelaphinae
Genus:Connochaetes
Hinrich Lichtenstein, 1812
Type species
Antilope gnu[1]
Species

Connochaetes gnou – black wildebeest
Connochaetes taurinus – blue wildebeest

Range map: 
Black wildebeest shown yellow
Blue wildebeest shown blue
Overlapping range shown brown

Wildebeest (/ˈwɪldɪbst/ WIL-dib-eest,[2][3][4] /ˈvɪl-/ VIL-,[4] /-dəb-/ -⁠dəb-), also called gnu (/nj/ new or /n/noo),[5][6][7] are antelopes of the genus Connochaetes and native to Eastern and Southern Africa. They belong to the family Bovidae, which includes true antelopes, cattle, goats, sheep, and other even-toed horned ungulates. There are two species of wildebeest: the black wildebeest or white-tailed gnu (C. gnou), and the blue wildebeest or brindled gnu (C. taurinus).

Fossil records suggest these two species diverged about one million years ago, resulting in a northern and a southern species. The blue wildebeest remained in its original range and changed very little from the ancestral species, while the black wildebeest changed more as adaptation to its open grassland habitat in the south. The most obvious ways of telling the two species apart are the differences in their colouring and in the way their horns are oriented.[8]

In East Africa, the blue wildebeest is the most abundant big-game species; some populations perform an annual migration to new grazing grounds, but the black wildebeest is merely nomadic. Breeding in both takes place over a short period of time at the end of the rainy season and the calves are soon active and are able to move with the herd, a fact necessary for their survival. Nevertheless, some fall prey to large carnivores, especially the spotted hyena.

Wildebeest often graze in mixed herds with zebra, which gives heightened awareness of potential predators. They are also alert to the warning signals emitted by other animals such as baboons. Wildebeest are a tourist attraction but compete with domesticated livestock for pasture and are sometimes blamed by farmers for transferring diseases and parasites to their cattle.[citation needed] Illegal hunting does take place but the population trend is fairly stable. Wildebeest can also be found in national parks or on private land. The International Union for Conservation of Nature lists both kinds of wildebeest as least-concern species.

Etymology[edit]

Wildebeest is Dutch for 'wild beast', 'wild ox' or 'wild cattle' in Afrikaans (bees 'cattle'),[citation needed] The name was given by Dutch settlers who saw them on their way to the interior of South Africa in about 1700 because they resemble wild ox. The blue wildebeest was first known to westerners in the northern part of South Africa a century later, in the 1800s.[9]

Some sources claim the name gnu originates from the Khoekhoe name for these animals, t'gnu.[10] Others contend the name and its pronunciation in English go back to the word !nu: used for the black wildebeest by the San people.[11][additional citation(s) needed]

Classification[edit]

Taxonomy and evolution[edit]

The genus name Connochaetes derives from the Ancient Greek words κόννοςkónnos 'beard',[12] and χαίτηkhaítē 'flowing hair, mane'.[13][14] It is placed under the family Bovidae and subfamily Alcelaphinae, where its closest relatives are the hartebeest (Alcelaphus spp.), the hirola (Beatragus hunteri), and species in the genus Damaliscus, such as the topi, the tsessebe, the blesbok and the bontebok.[15] The name Connochaetes was given by German zoologist Hinrich Lichtenstein in 1812.[15][16]

In the early 20th century, one species of the wildebeest, C. albojubatus, was identified in eastern Africa. In 1914, two separate races of the wildebeest were introduced, namely Gorgon a. albojubatus ("Athi white-bearded wildebeest") and G. a. mearnsi ("Loita white-bearded wildebeest"). However, in 1939, the two were once again merged into a single race, Connochaetes taurinus albojubatus. In the mid-20th century, two separate forms were recognised, Gorgon taurinus hecki and G. t. albojubatus.[9] Finally, two distinct types of wildebeest – the blue and black wildebeest – were identified. The blue wildebeest was at first placed under a separate genus, Gorgon,[17][18] while the black wildebeest belonged to the genus Connochaetes. Today, they are united in the single genus Connochaetes, with the black wildebeest being named (C. gnou) and the blue wildebeest (C. taurinus).[15]

According to a mitochondrial DNA analysis, the black wildebeest are estimated to have diverged from the main lineage during the Middle Pleistocene and became a distinct species around a million years ago.[19] A divergence rate around 2% has been calculated.[18] The split does not seem to have been driven by competitionfor resources, but instead because each species adopted a different ecological niche and occupied a different trophic level.[20][failed verification]

Blue wildebeest fossils dating back some 2.5 million years ago are common and widespread. They have been found in the fossil-bearing caves at the Cradle of Humankind north of Johannesburg. Elsewhere in South Africa, they are plentiful at such sites as ElandsfonteinCornelia, and Florisbad.[21] The earliest fossils of the black wildebeest were found in sedimentary rock in Cornelia in the Free State and dated back about 800,000 years.[20] Today, five subspecies of the blue wildebeest are recognised, while the black wildebeest has no named subspecies.[22][23]

Genetics and hybrids[edit]

The diploid number of chromosomes in the wildebeest is 58.[24] Chromosomes were studied in a male and a female wildebeest. In the female, all except a pair of very large submetacentric chromosomes were found to be acrocentricMetaphases were studied in the male's chromosomes, and very large submetacentric chromosomes were found there, as well, similar to those in the female both in size and morphology. Other chromosomes were acrocentric. The X chromosome is a large acrocentric and the Y chromosome a minute one.[16][25]

The two species of the wildebeest are known to hybridise. Male black wildebeest have been reported to mate with female blue wildebeest and vice versa.[26] The differences in social behaviour and habitats have historically prevented interspecific hybridisation between the species, but hybridisation may occur when they are both confined within the same area. The resulting offspring are usually fertile. A study of these hybrid animals at Spioenkop Dam Nature Reserve in South Africa revealed that many had disadvantageous abnormalities relating to their teeth, horns, and the wormian bones in the skull.[27] Another study reported an increase in the size of the hybrid as compared to either of its parents. In some animals, the tympanic part of the temporal bone is highly deformed, and in others, the radius and ulna are fused.[28]

Characteristics of the species[edit]

Wildebeest
Blue wildebeest
Black wildebeest

Both species of wildebeest are even-toed, horned, greyish-brown ungulates resembling cattle. Males are larger than females and both have heavy forequarters compared to their hindquarters. They have broad muzzles, Roman noses, and shaggy manes and tails.[29] The most striking morphological differences between the black and blue wildebeest are the orientation and curvature of their horns and the colour of their coats. The blue wildebeest is the bigger of the two species. In males, blue wildebeest stand 150 cm (59 in) tall at the shoulder and weigh around 250 kg (550 lb), while the black wildebeest stands 111–120 cm (44–47 in) tall[30] and weighs about 180 kg (400 lb). In females, blue wildebeest have a shoulder height of 135 cm (53 in) and weigh 180 kg (400 lb) while black wildebeest females stand 108 cm (43 in) at the shoulder and weigh 155 kg (342 lb). The horns of blue wildebeest protrude to the side, then curve downwards before curving up back towards the skull, while the horns of the black wildebeest curve forward then downward before curving upwards at the tips. Blue wildebeest tend to be a dark grey colour with stripes, but may have a bluish sheen. The black wildebeest has brown-coloured hair, with a mane that ranges in colour from cream to black, and a cream-coloured tail. The blue wildebeest lives in a wide variety of habitats, including woodlands and grasslands, while the black wildebeest tends to reside exclusively in open grassland areas.[22] In some areas, the blue wildebeest migrates over long distances in the winter, whereas the black wildebeest does not.[31] The milk of the black wildebeest contains a higher protein, lower fat, and lower lactose content than the milk of the blue wildebeest.[32] Wildebeest can live more than 40 years, though their average lifespan is around 20 years.[33] [34]

Distribution and habitat[edit]

Wildebeest inhabit the plains and open woodlands of parts of Africa south of the Sahara. The black wildebeest is native to the southernmost parts of the continent.[35] Its historical range included South Africa, Eswatini, and Lesotho, but in the latter two countries, it was hunted to extinction in the 19th century. It has now been reintroduced to them and also introduced to Namibia, where it has become well established.[36] It inhabits open plains, grasslands, and Karoo shrublands in both steep mountainous regions and lower undulating hills at altitudes varying between 1,350 and 2,150 m (4,430 and 7,050 ft).[37] In the past, it inhabited the highveld temperate grasslands during the dry winter season and the arid Karoo region during the rains. However, as a result of widespread hunting, the black wildebeest no longer occupies its historical range or makes migrations, and is now largely limited to game farms and protected reserves.[38]

The blue wildebeest is native to eastern and southern Africa. Its range includes KenyaTanzaniaBotswanaZambiaZimbabweMozambiqueSouth AfricaEswatini and Angola.[39] It is no longer found in Malawi but has been successfully reintroduced into Namibia. Blue wildebeest are mainly found in short grass plains bordering bush-covered acacia savannas, thriving in areas that are neither too wet nor too dry. They can be found in habitats that vary from overgrazed areas with dense bush to open woodland floodplains.[37] In East Africa, the blue wildebeest is the most abundant big game species, both in population and biomass.[29] It is a notable feature of the Serengeti National Park in Tanzania, the Maasai Mara National Reserve in Kenya and the Liuwa Plain National Park in Zambia.[33]

Migration[edit]

Migration in wildebeest
Wildebeest herding and following zebra in the Serengeti National Park
Wildebeest in Masai Mara during the Great Migration
Wildebeest leaping into the Mara River during their Great Migration

Not all wildebeest are migratory. Black wildebeest herds are often nomadic or may have a regular home range of 1 km2(0.39 sq mi). Bulls may occupy territories, usually about 100 to 400 m (300 to 1,300 ft) apart, but this spacing varies according to the quality of the habitat. In favourable conditions, they may be as close as 9 m (30 ft), or they may be as far apart as 1,600 m (5,200 ft) in poor habitat.[17] Female herds have home ranges of about 250 acres (100 ha; 0.39 sq mi) in size. Herds of nonterritorial bachelor males roam at will and do not seem to have any restrictions on where they wander.[40]

Blue wildebeest have both migratory and sedentary populations. In the Ngorongoro, most animals are sedentary and males maintain a network of territories throughout the year, though breeding is seasonal in nature. Females and young form groups of about 10 individuals or join in larger aggregations, and nonterritorial males form bachelor groups.[41] In the Serengeti and Tarangire ecosystems, populations are mostly migratory, with herds consisting of both sexes frequently moving, but resident subpopulations also exist.[42] During the rutting season, the males may form temporary territories for a few hours or a day or so, and attempt to gather together a few females with which to mate, but soon they have to move on, often moving ahead to set up another temporary territory.[41]

In the Maasai Mara game reserve, a non-migratory population of blue wildebeest had dwindled from about 119,000 animals in 1977 to about 22,000 in 1997. The reason for the decline is thought to be the increasing competition between cattle and wildebeest for a diminishing area of grazing land as a result of changes in agricultural practices, and possibly fluctuations in rainfall.[43]

Each year, some East African populations of blue wildebeest have a long-distance migration, seemingly timed to coincide with the annual pattern of rainfall and grass growth.[44] The timing of their migrations in both the rainy and dry seasons can vary considerably (by months) from year to year. At the end of the wet season (May or June in East Africa), wildebeest migrate to dry-season areas in response to a lack of surface (drinking) water. When the rainy season begins again (months later), animals quickly move back to their wet-season ranges. Factors suspected to affect migration include food abundance, surface water availability, predators, and phosphorus content in grasses. Phosphorus is a crucial element for all life forms, particularly for lactating female bovids. As a result, during the rainy season, wildebeest select grazing areas that contain particularly high phosphorus levels.[29] One study found, in addition to phosphorus, wildebeest select ranges containing grass with relatively high nitrogen content.[45]

Aerial photography has revealed that a level of organisation occurs in the movement of the herd that cannot be apparent to each individual animal; for example, the migratory herd exhibits a wavy front, and this suggests that some degree of local decision-making is taking place.[46] Numerous documentaries feature wildebeest crossing rivers, with many being eaten by crocodiles or drowning in the attempt. While having the appearance of a frenzy, recent research has shown a herd of wildebeest possesses what is known as a "swarm intelligence", whereby the animals systematically explore and overcome the obstacle as one.[47]

Interdigital glands[edit]

The wildebeest only has interdigital (hoof) glands in its fore legs. Analysis of chemical constituents from a free ranging blue wildebeest from Zimbabwe (Cawston Block) showed this gland contains cyclohexanecarboxylic acid, phenol, 2-phenolethanol, and six short-chain carboxylic acids. .[48]

Ecology[edit]

Predators[edit]

Taxidermied Lion and Blue WildebeestNamibia

Major predators that feed on wildebeest include the lionhyenaAfrican wild dogcheetahleopard, and Nile crocodile, which seem to favour the wildebeest over other prey.[33] Wildebeest, however, are very strong, and can inflict considerable injury even to a lion. Wildebeest have a maximum running speed of around 80 km/h (50 mph).[49][50] The primary defensive tactic is herding, where the young animals are protected by the older, larger ones, while the herd runs as a group. Typically, the predators attempt to isolate a young or ill animal and attack without having to worry about the herd. Wildebeest have developed additional sophisticated cooperative behaviours, such as animals taking turns sleeping while others stand guard against a night attack by invading predators. Wildebeest migrations are closely followed by vultures, as wildebeest carcasses are an important source of food for these scavengers. The vultures consume about 70% of the wildebeest carcasses available. Decreases in the number of migrating wildebeest have also had a negative effect on the vultures.[51] In the Serengeti ecosystem, Tanzania, wildebeest may help facilitate the migration of other, smaller-bodied grazers, such as Thomson's gazelles (Eudorcas thomsonii), which eat the new-growth grasses stimulated by wildebeest foraging.[52]

Interactions with nonpredators[edit]

Zebras and wildebeest group together in open savannah environments with high chances of predation. This grouping strategy reduces predation risk because larger groups decrease each individual's chance of being hunted, and predators are more easily seen in open areas.[53] The seasonal presence of thousands of migratory wildebeests reduces local lion predation on giraffe calves, resulting in greater survival of giraffes.[54]

Wildebeest can also listen in on the alarm calls of other species, and by doing so, can reduce their risk of predation. One study showed, along with other ungulates, wildebeests responded more strongly to the baboon alarm calls compared to the baboon contest calls, though both types of calls had similar patterns, amplitudes, and durations. The alarm calls were a response of the baboons to lions, and the contest calls were recorded when a dispute between two males occurred.[55] Wildebeest compete with domesticated livestock for pasture and are sometimes blamed by farmers for transferring diseases and parasites to their cattle.[56]

Breeding and reproduction[edit]

Video of wildebeest feeding its calf

Wildebeest do not form permanent pair bonds and during the mating season, or rut, the males establish temporary territories and try to attract females into them. These small territories are about 3,000 square metres (32,000 sq ft; 0.74 acres), with up to 300 territories per 1 square kilometre (0.39 sq mi; 250 acres). The males defend these territories from other males while competing for females that are coming into oestrus. The males use grunts and distinctive behaviour to entice females into their territories. Wildebeest usually breed at the end of the rainy season when the animals are well fed and at their peak of fitness.[29] This usually occurs between May and July, and birthing usually takes place between January and March, at the start of the wet season. Wildebeest females breed seasonally and ovulate spontaneously.[57]

The estrous cycle is about 23 days and the gestation period lasts 250 to 260 days. The calves weigh about 21 kg (46 lb) at birth[14] and scramble to their feet within minutes, being able to move with the herd soon afterwards,[33] a fact on which their survival relies.[58] The main predator of the calves is the spotted hyena. The calving peak period lasts for 2–3 weeks, and in small subpopulations and isolated groups, mortality of calves may be as high as 50%. However, in larger aggregations, or small groups living near large herds, mortality rates may be under 20%.[59]

Groups of wildebeest females and young live in the small areas established by the male. When groups of wildebeest join, the female to male ratio is higher because the females choose to move to the areas held by a smaller number of males.[53] This female-dominated sex ratio may be due to illegal hunting and human disturbance, with higher male mortality having been attributed to hunting.[60]

Threats and conservation[edit]

The black wildebeest has been classified as a least-concern species by the International Union for Conservation of Nature in the IUCN Red List. The populations of this species are on an increase. Now, more than 18,000 individuals are believed to remain, 7,000 of which are in Namibia, outside their natural range, and where it is farmed. Around 80% of the wildebeest occurs in private areas, while the other 20% is confined in protected areas. Its introduction into Namibia has been a success and numbers have increased substantially there from 150 in 1982 to 7,000 in 1992.[36]

Bag made with wildebeest skin

The blue wildebeest has also been rated as of least concern. The population trend is stable, and their numbers are estimated to be around 1,500,000 – mainly due to the increase of the populations in Serengeti National Park(Tanzania) to 1,300,000 as of 1998.[61] However, the numbers of one of the subspecies, the eastern white-bearded wildebeest (C. t. albojubatus) have seen a steep decline.[42] Population density ranges from 0.15/km2. in Hwange and Etosha National Parks to 35/km2 in Ngorongoro Conservation Area and Serengeti National Park.[37]

Overland migration as a biological process requires large, connected landscapes, which are increasingly difficult to maintain, particularly over the long term, when human demands on the landscape compete. The most acute threat comes from migration barriers, such as fences and roads. In one of the more striking examples of the consequences of fence-building on terrestrial migrations, Botswanan authorities placed thousands of kilometres of fences across the Kalahari that prevented wildebeests from reaching watering holes and grazing grounds, resulting in the deaths of tens of thousands of individuals, reducing the wildebeest population to less than 10% of its previous size.[62] Illegal hunting is a major conservation concern in many areas, along with natural threats posed by main predators (which include lions, leopards, African hunting dogs, cheetahs and hyenas). Where the black and blue wildebeest share a common range, the two can hybridise, and this is regarded as a potential threat to the black wildebeest.[26]

Uses and interaction with humans[edit]

Wildebeest provide several useful animal products. The hide makes good-quality leather and the flesh is coarse, dry and rather hard.[17] Wildebeest are killed for food, especially to make biltong in Southern Africa. This dried game meat is a delicacy and an important food item in Africa.[29] The meat of females is more tender than that of males, and is the most tender during the autumn season.[31] Wildebeest are a regular target for illegal meat hunters because their numbers make them easy to find. Cooks preparing the wildebeest carcass usually cut it into 11 pieces. The estimated price for wildebeest meat was about US$0.47 per 1 kilogram (2.2 lb) around 2008.[63] The silky, flowing tail of the black wildebeest is used to make fly-whisks or chowries.[17]

Wildebeest benefit the ecosystem by increasing soil fertility with their excreta. They are economically important for human beings, as they are a major tourist attraction. They also provide important products, such as leather, to humans.[64] Wildebeest, however, can also have a negative impact on humans. Wild individuals can be competitors of commercial livestock, and can transmit diseases and cause epidemics among animals, particularly domestic cattle. They can also spread tickslungwormstapeworms, flies, and paramphistome flukes.[9]

Cultural depictions[edit]

The wildebeest is depicted on the coat of arms of the Province of Natal and later KwaZulu-Natal in South Africa. Over the years, the South African authorities have issued several stamps displaying the animal and the South African Mint has struck a two-cent piece with a prancing black wildebeest.[65]

Movies and television shows also feature wildebeests, including Khumba (Mama V), The Wild (Kazar and his minions), All Hail King Julien (Vigman Wildebeest), Phineas and Ferb (Newton the Gnu), The Great Space Coaster (newscaster Gary Gnu), and The Lion King (the wildebeest stampede that resulted in Mufasa's death).

Michael Flanders wrote a humorous song called "The Gnu", which was very popular when he performed it, with Donald Swann, in a revue called At the Drop of a Hat, which opened in London on 31 December 1956.

In the 1970s, British tea brand Typhoo ran a series of television advertisements featuring an animated anthropomorphic gnu character.

The wildebeest is the mascot of the free software project GNU.

The wildebeest is the mascot of the GNU Project[66] and GNU operating system.

In the Llama Llama picture-book series by Anna Dewdney, an anthropomorphised wildebeest named Nelly Gnu is the main character, Llama Llama's best friend, and is also featured in a title of her own, Nelly Gnu and Daddy Too.[67]

In Terry Pratchett's Discworld series, Discworld analogues of wildebeests, which inhabited Howonderland, were called "bewilderbeest".

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